Novae Gesneriaceae neotropicarum XIII: Four new species of Columnea (Gesneriaceae) section Collandra from Colombia

Four new species of Columnea (Gesneriaceae) section Collandra (syn. Dalbergaria Tussac) are described from the remnant forests of the western and central cordilleras of Colombia: C. coronocrypta, C. coronata, C. pedunculata, and C. queremalensis. The flowers of C. coronata are unusual in the genus in having a corona, as in genera of Gesneriaceae pollinated by euglossine bees. Columnea pedunculata is distinct in having pedunculate inflorescences.